Modequai is the father of Morella. The two of them came to Greenwich ten years after the events of The Final Reckoning. Their appearance was quite a shock, as black squirrels were supposedly a dead race. There were rumors that Modequai and his daughter were the last surviving members of the Hawthorn Realm.

Soon after their arrival, Modequai barged into the chamber of the Starwife, Audrey Scuttle, and tore the silver acorn from about her neck. Then his daughter became the new Starwife as the 'Usurping Mouse' was chased away.